Huge crowds lined the streets of the holy city of Mashhad for the burial of Iran’s late supreme leader, Ayatollah Ali Khamenei.

Khamenei was laid to rest at the shrine of Imam Reza, Iran’s holiest Shia Muslim site, marking the end of six days of public mourning ceremonies in five cities across Iran and neighbouring Iraq.

The burial, reported by the state news agency IRNA, came after an exchange of strikes between Iran and the US that threatens to derail a preliminary deal to end the war in which he was killed.

Earlier, Iran’s Revolutionary Guards accused the US of bombing two bridges on the railway line from the capital, Tehran, to Mashhad overnight “in an effort to overshadow” the funeral.

Khamenei and several members of his family were killed in an Israeli strike on his residence in Tehran on 28 February, the first day of Iran’s war with the US and Israel.

He was succeeded as supreme leader by his son Mojtaba, who has not been seen in public since he was reportedly seriously wounded in the same attack.

The 56-year-old did not participate in the funeral ceremonies in Tehran and Qom, and there was no indication that he attended the burial.

On Thursday morning, a plane carrying the coffins of Ali Khamenei, his granddaughter, son-in-law, daughter and Mojtaba’s wife landed in Mashhad after flying from Iraq, where huge crowds took part in processions to two Shia shrines in the cities of Najaf and Karbala.

In the afternoon, Iranian TV footage showed thousands of mourners dressed in black walking along a main boulevard in central Mashhad. Many were waving Iranian flags and red banners symbolising vengeance.

Some people were also holding photos of the late supreme leader and placards calling for the death of US President Donald Trump, who, along with Israeli Prime Minister Benjamin Netanyahu, ordered the joint attack on Iran four months ago that triggered the war.

Hanging above the boulevard were banners with official slogans including “We must rise”.

“The loss of the leader is heavier than losing our parents,” Hoda, a 35-year-old housewife, told AFP news agency. “Only the death of Trump and Netanyahu will soothe our pain.”

Later, a lorry transported Khamenei’s coffin slowly through the crowds towards the Imam Reza shrine, eventually reaching the complex as night fell.

Imam Reza was the eighth Shia imam and the only one of the 12 believed to have been buried in Iran. His mausoleum, which dates to the 9th Century and has a towering golden dome and minarets, is visited by millions of pilgrims each year.